The Checklist of the Spiders of Iran (Arachnida: Araneae) --- Version 2021

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Araneidae Clerck, 1757

1)   Aculepeira armida (Audouin, 1826)

Records: Hamedan.

Distribution: North Africa, Southern Europe, Turkey, Israel, Russia (Europe to Far East), Iran, Central Asia to China.

References:

A. a.: Zamani et al., 2019b: 234.

2)   Aculepeira ceropegia (Walckenaer, 1802)

Records: Kerman.

Distribution: Europe, Turkey, Caucasus, Russia (Europe to West Siberia), Kazakhstan, Iran (?).

References:

Araneus ceropegius: Roewer, 1955: 754.

Comment: This species is probably misidentified and may refer to A. armida (Audouin, 1826).

3)   Aculepeira talishia (Zawadsky, 1902)

Records: Alborz, Kerman, Mazandaran, West Azerbaijan, Zanjan.

Distribution: Turkey, Iran, Caucasus to Central Asia.

References:

A. t.: Zamani et al., 2015: 340.

A. t.: Zamani & Mozaffarian, 2017: 8.

A. t.: Zamani et al., 2017b: 59.

4)   Agalenatea redii (Scopoli, 1763)

Records: Golestan, Kurdistan, Mazandaran, Tehran, Zanjan.

Distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia, China.

References:

A. r.: Ghavami, 2006: 1839.

A. r.: Ghahari & Marusik, 2009: 1.

A. r.: Ghahari & Tabari, 2012: 138.

A. r.: Kashefi et al., 2013: 9.

A. r.: Zamani, 2015: 13.

A. r.: Zamani et al., 2018a: 189.

5)   Araneus angulatus Clerck, 1757

Records: Gilan, Golestan, Hamedan, Isfahan, Kermanshah, Mazandaran, Qazvin, Razavi Khorasan, Semnan, Tehran.

Distribution: Europe, Turkey, Russia (Europe to Far East), Iran, Central Asia, Korea.

References:

A. a.: Mozaffarian, Tirgari & Bayat-Asadi, 2000: 65.

A. a.: Ghavami, 2006: 1839.

A. a.: Ghahari & Marusik, 2009: 2.

A. agulatus: Ghahari & Tabari, 2012: 138.

A. a.: Kashefi et al., 2013: 9.

A. a.: Zamani, 2015: 13.

A. a.: Zamani, 2016b: 132.

A. a.: Zamani et al., 2017b: 59.

A. a.: Zamani et al., 2019b: 234.

A. a.: Zamani et al., 2020: 570.

6)   Araneus circe (Audouin, 1826)

Records: Tehran, Zanjan.

Distribution: Southern Europe, Egypt, Turkey, Caucasus, Iran.

References:

A. c.: Zamani et al., 2015: 340.

A. c.: Zamani, 2016b: 133.

7)   Araneus diadematus Clerck, 1757

Records: Ardabil, East Azerbaijan, Gilan, Golestan, Mazandaran, Qazvin, Tehran.

Distribution: Europe, Middle East, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Japan. Introduced to North America.

References:

A. d.: Ghavami, 2006: 1840.

A. d.: Ghavami & Amooz, 2008: 74.

A. d.: Ghahari & Marusik, 2009: 2.

A. d.: Nikdel, 2015: 33.

A. d.: Zamani, 2016b: 134.

A. d.: Zamani et al., 2019b: 234.

8)   Araneus grossus (C. L. Koch, 1844)

Records: Kurdistan.

Distribution: Europe to Central Asia.

References:

A. g.: Zamani & Andr, 2017: 6.

9)   Araneus marmoreus Clerck, 1757

Records: Golestan, Kerman, Mazandaran, South Khorasan, Tehran.

Distribution: North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Korea, Japan.

References:

A. m.: Kashefi et al., 2013: 9.

A. m.: Zamani, 2016b: 135.

A. m.: Zamani et al., 2017b: 59.

10)                      Araniella alpica (L. Koch, 1869)

Records: Golestan.

Distribution: Europe, Turkey, Caucasus, Russia (Europe to Western Siberia), Iran (?).

References:

A. a.: Ghavami, 2006: 1839.

Comment: This species is probably misidentified.

11)                      Araniella cucurbitina (Clerck, 1757)

Records: Gilan, Golestan, Mazandaran, Tehran.

Distribution: Europe, Turkey, Russia (Europe) to Central Asia, China, Korea.

References:

Araneus cucurbitinus: Roewer, 1955: 754.

A. c.: Ghavami, 2006: 1839.

A. c.: Ghavami & Amooz, 2008: 74.

A. c.: Kashefi et al., 2013: 9.

A. c.: Zamani, 2016b: 136.

12)                      Araniella inconspicua (Simon, 1874)

Records: Golestan.

Distribution: Europe, Turkey, Russia (Europe to Far East), Iran (?), Central Asia to China.

References:

A. i.: Ghavami, 2006: 1839.

Comment: This species is probably misidentified.

13)                      Araniella mithra Zamani, Marusik & Šestáková, 2020*

Records: Chaharmahal & Bakhtiari, Isfahan, West Azerbaijan.

Distribution: Iran.

References:

A. proxima: Zamani et al., 2017b: 58 (misidentified per Zamani, Marusik & Šestáková, 2020: 15).

A. m. Zamani, Marusik & Šestáková, 2020: 15, f. 1AC, 2AB, 4AB, 6AB, 7C, 8C, 9C, 10C (♂♀).

14)                      Araniella opisthographa (Kulczyński, 1905)

Records: Ardabil, Golestan, Mazandaran.

Distribution: Europe, Turkey, Caucasus, Russia (Europe) to Central Asia, Iran.

References:

A. o.: Ghavami, 2006: 1839.

A. o.: Zamani et al., 2018a: 189.

A. o.: Zamani et al., 2019b: 234.

A. o.: Zamani, Marusik & Šestáková, 2020: 23, f. 2CD, 6CD, 7D, 8D, 9D, 10D (♂♀).

15)                      Araniella villanii Zamani, Marusik & Šestáková, 2020

Records: Chaharmahal & Bakhtiari.

Distribution: Iran, Kazakhstan, India.

References:

A. v. Zamani, Marusik & Šestáková, 2020: 18, f. 1B, D, 3AB, 4CD, 5AB, 7A, 8A, 9A, 10A (♂♀).

16)                      Argiope ahngeri Spassky, 1932

Records: Alborz.

Distribution: Iran, Kyrgyzstan, Turkmenistan, Uzbekistan, Tajikistan (?).

References:

A. a.: Zamani et al., 2018a: 184.

17)                      Argiope anasuja Thorell, 1887

Records: Sistan & Baluchistan.

Distribution: Seychelles, Maldives, Iran, Pakistan, India, Sri Lanka, Australia (Cocos Is.).

References:

A. a.: Zamani et al., 2017b: 58.

18)                      Argiope bruennichi (Scopoli, 1772)

Records: Chaharmahal & Bakhtiari, East or West Azerbaijan, Fars, Gilan, Golestan, Mazandaran, Qazvin, Razavi Khorasan, Zanjan.

Distribution: Europe, Turkey, Israel, Russia (Europe to Far East), Iran, Central Asia to China, Korea, Japan.

References:

A. b.: Pocock, 1899: 400.

Argyope brnnichii: Roewer, 1955: 754 (lapsus).

A. b.: Mozaffarian & Tirgari, 2000: 104.

A. b.: Mozaffarian, Tirgari & Bayat-Asadi, 2000: 65.

A. b.: Kakhki, 2005b: 64.

A. b.: Mirshamsi & Darvish, 2005: 408, f. 69 (♂♀).

A. b.: Ghavami et al., 2007: 2564.

A. b.: Ghavami & Amooz, 2008: 74.

A. b.: Ghahari & Marusik, 2009: 2.

A. b.: Ghahari & Tabari, 2012: 138.

A. b.: Jäger, 2012: 287.

A. b.: Kashefi et al., 2013: 9.

A. b.: Zamani, 2015: 13.

A. b.: Sadeghi et al., 2016: 5.

A. b.: Zamani & Mozaffarian, 2017: 8.

A. b.: Zamani et al., 2017b: 59.

19)                      Argiope lobata (Pallas, 1772)

Records: Ardabil, Fars, Golestan, Kerman, Mazandaran, Razavi Khorasan, South Khorasan, Tehran, Yazd, Zanjan.

Distribution: Southern Europe to Central Asia and China, northern Africa, South Africa, Israel, India, Myanmar to New Caledonia and northern Australia.

References:

A. l.: Pocock, 1889: 110.

Argyope l.: Roewer, 1955: 754 (lapsus).

A. l.: Kakhki & Darvish, 2002: 162, f. 6, 9a, 10 (♂♀).

A. l.: Kakhki, 2005b: 64.

A. l.: Mirshamsi & Darvish, 2005: 406, f. 15 (♂♀).

A. l.: Ghavami, 2006: 1840.

A. l.: Ghavami, 2008: 687.

A. l.: Ghahari & Tabari, 2012: 139.

A. l.: Jäger, 2012: 289.

A. l.: Kashefi et al., 2013: 9.

A. l.: Namaghi, Kaykhosravi & Zamani, 2014: 106.

A. l.: Zamani, 2015: 13.

A. l.: Zamani & Mozaffarian, 2017: 9.

A. l.: Zamani et al., 2019b: 234.

20)                      Argiope sector (Forsskål, 1776)

Records: Bushehr, Hormozgan.

Distribution: Cape Verde Is., Senegal, North Africa, Middle East.

References:

A. clarki: Pocock, 1903: 188, pl. 14, f. 3 (♂♀).

A. s.: Zamani et al., 2017b: 59.

21)                      Argiope trifasciata (Forsskål, 1775)

Records: Hormozgan.

Distribution: North, Central and South America. Introduced to Africa, Portugal to Israel, Iran, China, Japan, Australia (Tasmania), Pacific Is.

References:

A. t.: Zamani et al., 2014b: 30.

22)                      Cyclosa conica (Pallas, 1772)

Records: Gilan, Golestan, Mazandaran, Qazvin, Zanjan.

Distribution: North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China.

References:

C. c.: Ghavami, 2006: 1840.

C. c.: Ghavami et al., 2007: 2564.

C. c.: Ghavami & Amooz, 2008: 74.

C. c.: Ghahari & Tabari, 2012: 139.

C. c.: Zamani, 2015: 13.

C. c.: Zamani et al., 2019b: 234, f. 1AD (♂♀).

23)                      Cyclosa sierrae Simon, 1870

Records: Golestan, Hormozgan, Kohgiluyeh & Boyer-Ahmad, Mazandaran.

Distribution: Southern Europe, Hungary, Ukraine, Turkey, Caucasus, Iran.

References:

C. s.: Kashefi et al., 2013: 9.

C. s.: Zamani et al., 2019b: 234.

24)                      Cyrtophora citricola (Forsskål, 1775)

Records: Hormozgan, Kohgiluyeh & Boyer-Ahmad.

Distribution: Southern Europe, Africa, Middle East, Pakistan, India, China, Japan. Introduced to Dominican Rep., Costa Rica, Colombia, Brazil.

References:

C. cf. c.: Zamani et al., 2017b: 58.

C. c.: Hosseinpour et al., 2019: 206.

25)                      Gibbaranea bituberculata (Walckenaer, 1802)

Records: Fars, Isfahan.

Distribution: North Africa, Europe, Turkey, Israel, Russia (Europe to Far East), Iran, Central Asia to China, Japan, India.

References:

G. b.: Zamani et al., 2019b: 234.

26)                      Gibbaranea ullrichi (Hahn, 1835)

Records: Chaharmahal & Bakhtiari.

Distribution: Europe to Central Asia.

References:

G. u.: Zamani et al., 2019b: 236.

27)                      Hypsosinga albovittata (Westring, 1851)

Records: Kermanshah, Tehran, West Azerbaijan.

Distribution: Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Middle East, Central Asia.

References:

H. a.: Ghavami, 2006: 1840.

H. a.: Zamani et al., 2019b: 236.

H. a.: Zamani et al., 2020: 570.

28)                      Hypsosinga heri (Hahn, 1831)

Records: Fars.

Distribution: Europe, Caucasus, Israel, Russia (Europe to Central Asia, West Siberia), Israel, Iran, Central Asia, China.

References:

H. h.: Zamani et al., 2015: 340.

29)                      Hypsosinga pygmaea (Sundevall, 1831)

Records: Gilan, Golestan, Isfahan, Kermanshah, Mazandaran, North Khorasan (?), Razavi Khorasan (?), West Azerbaijan, Zanjan.

Distribution: North America, Europe, Turkey, Israel, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Korea, Japan.

References:

H. p.: Mozaffarian, Tirgari & Bayat-Asadi, 2000: 65.

H. p.: Ghavami, 2006: 1840.

H. p.: Ghavami et al., 2007: 2564.

H. p.: Khodashenas et al., 2012: 131.

H. p.: Zamani, 2015: 13.

H. p.: Zamani & Mozaffarian, 2017: 9.

H. p.: Zamani et al., 2017b: 57.

H. p.: Zamani et al., 2018a: 189.

H. p.: Zamani et al., 2019b: 236.

30)                      Hypsosinga sanguinea (C. L. Koch, 1844)

Records: Golestan, Mazandaran, North Khorasan (?), Razavi Khorasan (?), West Azerbaijan.

Distribution: Europe, North Africa, Turkey, Russia (Europe to Far East), Iran, Central Asia, China, Korea, Japan.

References:

H. s.: Ghavami, 2006: 1840.

H. s.: Ghavami & Amooz, 2008: 74.

H. sanguine: Khodashenas et al., 2012: 131 (lapsus).

H. s.: Zamani et al., 2020: 570.

31)                      Larinia chloris (Audouin, 1826)

Records: Lorestan.

Distribution: North and East Africa to Israel, Turkey, Iran, India, Sri Lanka, Bangladesh.

References:

L. c.: Mozaffarian, 2000: 108.

32)                      Larinioides cornutus (Clerck, 1757)

Records: Golestan, Kerman, Zanjan.

Distribution: North America, Europe, Turkey, Israel, Caucasus, Russia (Europe to Far East), Iran, China, Korea, Japan.

References:

Araneus c.: Roewer, 1955: 754.

L. c.: Ghavami, 2006: 1840.

L. c.: Ghavami & Amooz, 2008: 74.

L. c.: Kashefi et al., 2013: 10.

L. c.: Zamani & Mozaffarian, 2017: 9.

33)                      Larinioides ixobolus (Thorell, 1873)

Records: Unspecified province.

Distribution: Western Europe to Central Asia.

References:

L. i.: Roelofs-Ditters, 2001: 3.

34)                      Larinioides suspicax (O. Pickard-Cambridge, 1876)

Records: Alborz, Chaharmahal & Bakhtiari, Gilan, Golestan, Hamedan, Isfahan, Kerman, Kermanshah, Lorestan, Mazandaran, Razavi Khorasan, Tehran, West Azerbaijan.

Distribution: Europe, North Africa to Central Asia.

References:

L. folium: Mozaffarian & Tirgari, 2000: 104.

L. folium: Mozaffarian, Tirgari & Bayat-Asadi, 2000: 65.

L. folium: Ghavami, 2006: 1840.

L. folium: Ghahari & Marusik, 2009: 2.

L. folium: Ghahari & Tabari, 2012: 139.

L. folium: Šestáková, Marusik & Omelko, 2014: 78, f. 7AD, 8G, 9G, 15AD, 16AD, 25B, 31CD (♂♀).

L. s.: Zamani et al., 2018a: 189.

L. s.: Zamani et al., 2019b: 236.

35)                      Leviellus caspicus (Simon, 1889)

Records: Tehran, West Azerbaijan, Yazd.

Distribution: Kazakhstan, Kyrgyzstan, Uzbekistan, Turkmenistan, Iran.

References:

L. c.: Zamani et al., 2015: 340.

L. c.: Zamani & Mozaffarian, 2017: 9.

L. c.: Zamani et al., 2019b: 236.

36)                      Mangora acalypha (Walckenaer, 1802)

Records: Alborz, Chaharmahal & Bakhtiari, Gilan, Golestan, Isfahan, Kermanshah, Markazi, Mazandaran, North Khorasan (?), Qazvin, Razavi Khorasan (?), Semnan, Tehran.

Distribution: Madeira, Europe, North Africa, Turkey, Middle East, Caucasus, Russia (Europe to South Siberia), Central Asia, China.

References:

M. accalypha: Nazari, Mozaffarian & Marusik, 2002: 13 (lapsus).

M. a.: Ghavami, 2006: 1840.

M. a.: Ghavami et al., 2007: 2564.

M. a.: Ghavami, 2008: 687.

M. a.: Ghavami & Amooz, 2008: 74.

M. accalypha: Ghahari & Tabari, 2012: 139 (lapsus).

M. a.: Kashefi et al., 2013: 10.

M. a.: Zamani, 2016b: 144.

M. a.: Zamani & Mozaffarian, 2017: 9.

Mongora a.: Khodashenas et al., 2012: 131 (lapsus).

M. a.: Zamani et al., 2018a: 189.

M. a.: Zamani et al., 2019b: 236.

37)                      Neoscona adianta (Walckenaer, 1802)

Records: Alborz, Ardabil, Gilan, Golestan, Hamedan, Isfahan, Kermanshah, Kurdistan, Lorestan, Mazandaran, Qazvin, Tehran, West Azerbaijan, Zanjan.

Distribution: Europe, North Africa to Central Asia, Russia (Europe to Far East), China, Korea, Japan.

References:

N. a.: Mozaffarian & Tirgari, 2000: 103.

N. a.: Komposch, 2002: 103.

N. a.: Ghavami et al., 2007: 2564.

N. a.: Ghavami & Amooz, 2008: 74.

N. a.: Ghavami et al., 2008: 182.

N. a.: Ghahari & Marusik, 2009: 2.

N. a.: Ghahari & Tabari, 2012: 139.

N. a.: Kashefi et al., 2013: 10.

N. a.: Zamani, 2015: 13.

N. a.: Zamani, 2016b: 112.

N. a.: Zamani & Mozaffarian, 2017: 9.

N. a.: Zamani et al., 2018a: 189.

N. a.: Zamani et al., 2019b: 236.

38)                      Neoscona isatis Zamani, Marusik & Šestáková, 2020*

Records: Yazd.

Distribution: Iran.

References:

N. i. Zamani, Marusik & Šestáková, 2020: 28, f. 11DE, 13EF, 14B, 14F, 15EG, 16DF, 17GI (♂♀).

39)                      Neoscona spasskyi (Brignoli, 1983)

Records: Golestan, North Khorasan, Razavi Khorasan.

Distribution: Iran, Kazakhstan, Turkmenistan, Tajikistan, Kyrgyzstan.

References:

N. s.: Zamani, Marusik & Šestáková, 2020: 28, f. 11C, 12AB, 13CD, 14C, E, 15AC, 16AC, 17DF (♂♀).

40)                      Neoscona subfusca (C. L. Koch, 1837)

Records: Golestan, Hormozgan, Kerman, Markazi, Mazandaran, Qazvin, Yazd, Zanjan.

Distribution: Southern Europe, Africa, Turkey, Middle East, Ukraine, Caucasus, Russia (Europe) to Central Asia.

References:

N. s.: Ghahari & Marusik, 2009: 2.

N. s.: Ghahari & Tabari, 2012: 139.

N. s.: Kashefi et al., 2013: 10.

N. s.: Zamani, 2015: 13.

N. s.: Zamani & Mozaffarian, 2017: 9.

N. s.: Zamani et al., 2018a: 189.

N. s.: Zamani et al., 2019b: 236.

N. s.: Zamani et al., 2020: 570.

41)                      Neoscona theisi (Walckenaer, 1841)

Records: Gilan, Mazandaran.

Distribution: Cyprus, Israel, Caucasus (Georgia, Russia), Iran, Pakistan, India, Philippines, China to Indonesia, Japan, Australia. Introduced to Seychelles, Pacific Is.

References:

N. t.: Zamani, Marusik & Šestáková, 2020: 28, f. 11AB, 14A, D, 17AC (♂♀).

42)                      Nuctenea silvicultrix (C. L. Koch, 1835)

Records: Gilan.

Distribution: Europe, Russia (Europe to Far East), Kazakhstan.

References:

Araneus s.: Roewer, 1955: 754.

Comment: This species is probably misidentified.

43)                      Nuctenea umbratica (Clerck, 1757)

Records: Gilan, Golestan, Mazandaran.

Distribution: Europe to Iran.

References:

N. u.: Komposch, 2002: 103.

N. u.: Zamani, 2015: 13.

N. u.: Zamani et al., 2017b: 59.

N. u.: Zamani et al., 2019b: 236.

44)                      Poltys nagpurensis Tikader, 1982

Records: Hormozgan.

Distribution: Iran, India.

References:

P. n.: Zamani et al., 2019a: 4, f. 1af, 2ag, 3ah (♀).

45)                      Singa hamata (Clerck, 1757)

Records: Gilan, Mazandaran, Tehran, West Azerbaijan.

Distribution: Europe, Turkey, Russia (Europe to Far East), Caucasus to Central Asia, China, Korea, Japan.

References:

S. h.: Ghavami & Amooz, 2008: 74.

S. h.: Zamani & Mozaffarian, 2017: 9.

S. h.: Zamani et al., 2019b: 236.

46)                      Singa lucina (Audouin, 1826)

Records: Gilan, Golestan, Kerman, Kermanshah, Mazandaran, West Azerbaijan.

Distribution: Mediterranean to Central Asia.

References:

S. l.: Mozaffarian & Tirgari, 2000: 104.

S. l.: Mozaffarian, Tirgari & Bayat-Asadi, 2000: 65.

S. licina: Ghavami, 2006: 1840 (lapsus).

S. l.: Ghahari & Marusik, 2009: 2.

S. l.: Ghahari & Tabari, 2012: 139.

S. l.: Zamani, 2015: 13.

S. l.: Zamani et al., 2018a: 189.

S. l.: Zamani et al., 2019b: 236.

47)                      Singa semiatra L. Koch, 1867

Records: Golestan.

Distribution: Mediterranean, Ukraine, Russia (Europe), Iraq, Iran.

References:

S. s.: Kashefi et al., 2013: 10.

48)                      Siwa atomaria (O. Pickard-Cambridge, 1876)

Records: Kermanshah.

Distribution: Egypt, Israel, Azerbaijan, Iran.

References:

S. a.: Zamani et al., 2018a: 184.

49)                      Zilla diodia (Walckenaer, 1802)

Records: Isfahan, Mazandaran.

Distribution: North Africa, Europe, Turkey, Caucasus, Russia (Europe, West Siberia), Iran.

References:

Z. d.: Zamani et al., 2019b: 236.

50)                      Zygiella x-notata (Clerck, 1757)

Records: Golestan, North Khorasan (?), Razavi Khorasan (?).

Distribution: Europe, Turkey, Caucasus, Iran (?). Introduced to North America, Chile, Uruguay, Argentina, China, Japan, Runion.

References:

Z. x.: Ghavami, 2006: 1843.

Z. x.: Khodashenas et al., 2012: 131.

Comment: This species is probably misidentified.